Two new species of ant-loving beetles (Coleoptera: Staphylinidae: Pselaphinae) from Jiulong National Wetland Park, China.

Continued collecting efforts at the Jiulong National Wetland Park, Zhejiang, East China revealed two additional species of the ant-loving beetle subfamily Pselaphinae (Coleoptera: Staphylinidae): Trisiniotus jiulong sp. nov. and Arthromelodes lianghongbini sp. nov. Both new species are diagnosed, described, and their important characters are illustrated.

The genus Pseudophanias Raffray (Coleoptera, Staphylinidae, Pselaphinae) from Nanling Priority Area for Biodiversity Conservation, China.

Prior to this study, no species of Pseudophanias Raffray had been reported from Nanling, a vast biodiversity conservation area that spans five provinces in southern China. In this paper, three new species of the genus are described: Pseudophaniasfurcilobussp. nov. (Guizhou, Guangxi), P.leigongsp. nov. (Guizhou), and P.mulunsp. nov. (Guangxi), suggesting that additional study on the diversity of this group in the area is required. These species are characterized, keyed, and compared to similar congeners, supplemented with illustrations of the habitus and other morphological characters.

Redescription and a new record of Centrophthalmus sinensis Raffray, 1904(Coleoptera: Staphylinidae: Pselaphinae).

A new record of Centrophthalmus sinensis Raffray, 1904 from Suzhou, East China is provided, which is also the first Pselaphinae known to occur in this city, and the third pselaphine species from Jiangsu Province. A redescription of the species is given, aided with illustrations of the habitus and other morphological details.

A new species of Tyrodes Raffray (Coleoptera: Staphylinidae: Pselaphinae) from China.

A new species of the small pselaphine genus Tyrodes Raffray (Pselaphitae: Tyrini), T. tibialis sp. nov., is described and illustrated from Guangxi, southwestern China. This species can be separated from its congeners mainly by the aedeagal characters as well as by the modified sternite 2 (V), and presence of a preapical projection of the metatibiae.

A new species of Physomerinus Jeannel (Coleoptera, Staphylinidae, Pselaphinae) from Jiulong National Wetland Park, China.

Physomerinusjiulongensissp. nov. is described from a series of overwintering individuals collected in decomposing wood at Jiulong National Wetland Park, East China. The new species is characterized and separated from related congeners by the unique form of the sexually dimorphic maxillary palpi, greatly swollen male metafemora, as well as by the shape of the genitalia of both sexes. A key to, and a distributional map of, Physomerinus species occurring in China and on the Ryukyu Islands, Japan is provided.

A third species of Pakistatyrus Hlavác (Coleoptera: Staphylinidae: Pselaphinae).

The pselaphine genus Pakistatyrus Hlavác (Pselaphitae: Tyrini) is represented by two species distributed in high mountain areas of the Himalaya. Here a third species, P. schmidti sp. nov., is described from Nepal. The species is characterized by the simple male antennae, large foveae of the pronotum and elytra, greatly reduced elytra, and conspicuously asymmetric aedeagus. A key to and an updated distribution map of the species of Pakistatyrus is provided.

Range extension and male description of a rare clavigerite beetle in China (Coleoptera: Staphylinidae: Pselaphinae).

The myrmecophilous clavigerite Anaclasiger zhudaiae Yin & Huang is known from a single female collected in Guangdong, southern China. Here, I report the discovery of this species in Hunan, central China, associated with Nylanderia flaviabdominis (Wang) ants, which extends its range approximately 630 km northward from the type locality. The male is described for the first time, and the habitus and other major diagnostic characters are illustrated. Ex-situ photographs of living beetles and a map showing the distribution of Anaclasiger Raffray are provided.

New material of Euroleptochromus tuberculatus Yin & Cai (Coleoptera: Staphylinidae: Scydmaeninae) in Eocene Baltic amber.

The extinct members of the ant-like stone beetle supertribe Mastigitae are among the best studied groups of all fossil Scydmaeninae, and their evolutionary history can be traced from the latest Albian to the earliest Cenomanian through Miocene/Oligocene, with seven extinct genera being described (summarized in Jaoszyski, 2016; Jaoszyski & Perkovsky, 2016; Jaoszyski et al., 2018; Ross, 20192022 (list of Burmese amber taxa)).

Two new riparian Pselaphinae from Hubei, Central China (Coleoptera: taphylinidae).

The Pselaphinae fauna of Hubei Province in central China is extremely poorly documented. Here we describe two new species recently collected at a riverbank near Chongyangs county seat. Batrisiella riparia sp. nov. is characterized by the greatly swollen metafemora of the male, and Nipponobythus varicornis sp. nov. has a darkish-colored body, and uniquely modified male antennal clubs and metaventrite. Illustrations of the habitus and taxonomically informative characters of both species are provided for a reliable identification.

A new Cephennodes Reitter (Coleoptera: Staphylinidae: Scydmaeninae) of C. lustricollis group from East China.

The C. lustricollis group of the scydmaenine genus Cephennodes Reitter was previously represented by three species distributed in northern Vietnam and China (Hong Kong and Guizhou). Here we describe a fourth member, Cephennodes gusu sp. nov., collected from Qionglong Mountain, Suzhou, eastern China. The new species is characterized by its moderately large body size, the punctation of the head, pronotum and elytra, and the structure of the aedeagus. It is also the first record of the subfamily Scydmaeninae in Jiangsu Province.

A new species of Tmesiphorus LeConte from northern Myanmar (Coleoptera: Staphylinidae: Pselaphinae).

A new species of the pselaphine genus Tmesiphorus, T. shihongliangi sp. nov., is described and illustrated from Kachin State, North Myanmar, and is compared with and separated from allied congeners.

Schmidtinus annapurnensis, a new genus and species of Batrisitae from Nepal (Coleoptera: Staphylinidae: Pselaphinae).

Schmidtinus annapurnensis Yin Kurbatov gen. et sp. nov. of the pselaphine tribe Batrisini is described from the Annapurna mountains of Nepal, with the habitus and major diagnostic features illustrated. The morphological features that distinguish Schmidtinus gen. nov. from related batrisine genera are discussed.

The Earliest Corotocini (Insecta: Coleoptera: Staphylinidae) from Dominican Amber, with Remarks on Post-Imaginal Growth Influence on Termitophile Taxonomy.

Pareburniola dominicana Zilberman, Yin & Cai gen. et. sp. nov. is the very first fossil record of the tribe Corotocini, reported from Miocene Dominican Republic amber. The new species, which is based on a stenogastric individual, is described and illustrated and is included in the subtribe Corotocina due to the combination of a tarsal formula 4-4-4, an elongated gula, a developed labial palp, a reduced fourth palpomere, separated metacoxae and a glandular structure on the posterior region of the head. Since the taxon belongs to the physogastric tribe Corotocini, which presents post-imaginal growth, this phenomenon is herein discussed, and its current knowledge is used to understand the possible outcomes during morphological changes in the fossil species.

A new species of Trisiniotus Jeannel from Mao-shan, East China (Coleoptera, Staphylinidae, Pselaphinae).

The genus Trisiniotus Jeannel of the pselaphine tribe Batrisini comprises two species distributed in North India and southern Myanmar. Here, a third species, T.taoismus Feng & Yin sp. nov., is described from Mao-shan, Jiangsu Province, East China. The new species can be readily distinguished from both congeners by the unmodified male antennae.

The Batrisini of Tibet: unveiling an enigmatic ant-loving beetle diversity at Earths Third Pole (Coleoptera, Staphylinidae, Pselaphinae).

The taxonomic knowledge of the pselaphine tribe Batrisini (Coleoptera: Staphylinidae) from the Tibet Autonomous Region is summarized, with 80 species placed in 13 genera. Two genera and 68 species are described as new, aided with illustrations of habitus and diagnostic characters: Arthromelodes alpitorus sp. nov., A. angulatus sp. nov., A. aniqiao sp. nov., A. bicolor sp. nov., A. complexus sp. nov., A. cona sp. nov., A. crassicornis sp. nov., A. criniger sp. nov., A. cylindricus sp. nov., A. flosculus sp. nov., A. gyamda sp. nov., A. intricatus sp. nov., A. lage sp. nov., A. langjicuo sp. nov., A. latithorax sp. nov., A. lebus sp. nov., A. lotus sp. nov., A. markam sp. nov., A. monba sp. nov., A. nepaeformis sp. nov., A. nomurai sp. nov., A. songxiaobini sp. nov., A. speciosus sp. nov., A. torus sp. nov., A. zhangmu sp. nov., A. zhentangensis sp. nov., Batriscenaulax campestris sp. nov., Batrisiella acclivis sp. nov., Batrisiella beibeng sp. nov., Batrisiella bomea sp. nov., Batrisiella concisa sp. nov., Batrisiella curvitibialis sp. nov., Batrisiella ganma sp. nov., Batrisiella guttata sp. nov., Batrisiella zhujianqingi sp. nov., Batrisodes guoguotang sp. nov., Batrisodes hanmi sp. nov., Coryphomimus levigatus gen. et sp. nov., Coryphomodes budda sp. nov., Coryphomodes cephalicus sp. nov., Coryphomodes chenzhilini sp. nov., Gyrongita uniformis gen. et sp. nov., Sathytes aequalis sp. nov., S. asura sp. nov., S. caichenyangi sp. nov., S. dawai sp. nov., S. duojii sp. nov., S. laticornis sp. nov., S. pengzhongi sp. nov., S. proclivis sp. nov., S. pseudograndis sp. nov., S. sympatricus sp. nov., S. virupaksa sp. nov., S. yigongensis sp. nov., Tribasodites bari sp. nov., T. cellulanus sp. nov., T. constrictus sp. nov., T. dilophus sp. nov., T. elongatus sp. nov., T. grandiceps sp. nov., T. gyirong sp. nov., T. kiypu sp. nov., T. mirabilis sp. nov., T. prolixicornis sp. nov., T. pugiunculus sp. nov., T. vertexalis sp. nov., T. yatung sp. nov., and Trisinus quadrilobus sp. nov. The genus Plocamarthrus Jeannel, 1960 is proposed as a subjective junior synonym of Arthromelodes Jeannel 1954, syn. nov., resulting in Arthromelodes championi (Jeannel, 1960) comb. nov. (type species of Plocamarthrus), A. cameroni (Jeannel, 1960) comb. nov., A. planiceps (Jeannel, 1960) comb. nov., and A. abdominalis (Jeannel, 1960) comb. nov. Arthromelodes championi and Sathytes simplex Lbl, 1979 are reported to occur in Tibet, which are also new country records for China. Additional collecting data are provided for Hingstoniella lata Jeannel, 1960, Sathytes panzhaohuii Yin Shen, 2020, Sathytes transversus Yin Shen, 2020, and Sathytes xizangensis Yin Shen, 2020. Keys for identification of the genera and species are presented. Maps showing the distribution of each species are provided, supplemented with habitat photos. Lists of Tibetan species of Batrisini and of other pselaphine groups are given, adding up to 95 species.

Two New Species of Batrisini (Coleoptera: Staphylinidae: Pselaphinae) from Nanling Mountain Area, China.

Two new species of the Batrisocenus complex of genera, e.g., Batrisceniola nanlingensis sp. nov. and Physomerinus clavipes sp. nov., are described, diagnosed, and illustrated from the Nanling Mountain Area. Both represent a second species of the respective genus from the Chinese mainland.

An unusual species of Pseudophanias Raffray from West Tianmu Mountain, China (Coleoptera, Staphylinidae, Pselaphinae).

The genus Pseudophanias Raffray includes 19 species distributed in the Oriental Region. Here, the first species from continental China, P.tianmuensis Yin & Zhao sp. nov., is described and illustrated from West Tianmu Mountain. This species can be readily separated from all congeners by the coarse vestiture, along with its male features. A previous key to Pseudophanias species of East and South Asia is modified to accommodate the new species.